Afzal-Rafii, Z.. 1980: Contribution à ľ étude cytotaxonomique de quelques Cousinia d’Iran. -Biol. Ecol. Medit. 7: 6-14.
Aryavand, A. 1975: Contribution à ľ étude cytotaxonomique de quelques angiospermes de ľIran. -Botaniska Notiser 128: 299-311.
Assadi, M. 1987: Plan of the Flora of Iran (in Farsi). –Research Institute of Forests and Rangelands, Tehran.
Attar, F. 2000: The systematic studies of Cousinia (Asteraceae) sect. Cynaroideae in Iran (Ph.D. thesis). -Tehran University.
Attar, F. & Ghahreman, A. 2000: Two new species and a new record of the genus Cousinia Cass. Sect. Cynaroideae (Asteraceae) from west of Iran. -Iran Journ. Bot. 8 (2): 259-269.
Attar, F. & Ghahreman, A. 2005: Two new species of the genus Cousinia (Asteraceae) from Iran. -Nord. Journ. Bot. 23 (4): 589-592.
Attar, F. & Ghahreman, A. 2006: A synopsis of sect. Cynaroides (Cousinia, Compositae), distribution patterns and diversity centers. -Rostaniha 7 (Supplement 2): 315-342.
Attar, F. & Ghahreman, A. 2007: Cousinia aligudarzensis (Asteraceae), a new species of section Cynaroides Bunge from Iran. -Novon 17: 145-147.
Attar, F., Ghahreman, A. & Assadi, M. 2000: Three new species of the genus Cousinia (Asteraceae, Sect. Cynaroideae) from Iran. -Nord. Journ. Bot. 20 (6): 697-700.
Attar, F., Ghahreman, A. & Assadi, M. 2001: Studies on the genus Cousinia Cass. (Compositae) in Iran. -Iran Journ. Bot. 9 (1): 55-62.
Attar, F., Ghahreman, A. & Assadi, M. 2002: Cousinia zagrica (Asteraceae), a new species from West of Iran. -Sendtnera 8: 5-7.
Boisser, E., 1875: Cousinia in Flora Orientalis 3: 458-513. –Genevae and Basileae.
Bremer, K. 1994: Asteraceae: Cladistics and classification. -Timber press, Portland, Oregon.
Koch, C. 1851: Beiträge zu einer Flora des Orientes. -Linnaea 24:305-624.
Cassini, A. H. G. 1827: Cousinia Cass. in Dict. Sci. Nat. 47: 503. -Strasbourg.
DE Candolle, A. P. 1837: Cousinia in Prodromus Systematicis Naturalis Regni Vegetabilis 6: 552-557. –Parisiis.
Fedorov, AA. (ed.). 1969: Chromosome Numbers of Flowering Plants. -Leningrad: Nauka (in Russian).
Ghaffari, S. M. 1984: IOPB Chromosome number reports LXXXIII. -Taxon 33: 351-354.
Ghaffari, S. M. 1986: Chromosome number reports XCIII. -Taxon 35: 897-903.
Ghaffari, S. M. 1987: Chromosome studies in some flowering plants of Iran. -Rev. Cytol. Biol. Veg. Bot. 10: 3-8.
Ghaffari, S. M., Attar, F. & Ghahreman 2000 Distribution and chromosome studies on some species of Cousinia Cass., (section Cynaroideae) from Iran. -Pak. Journ. Bot. 32 (2): 311-316.
Ghaffari, S. M., Garcia-Jacas, N. & Susanna, A. 2006: New chromosome counts in the genus Cousinia (Asteraceae) from Iran. -The Linnean Society of London, Botanical Journal of the Linnean Society 151: 411-419.
Ghahreman, A., Iranshahr, M. & Attar, F. 1999: Introduction two new and a rare species of the genus Cousinia Cass., sect. Cynaroideae (Asteraceae). -Iran Journ. Bot. 8 (1): 15-22.
Haffner, E. 2000: On the phylogeny of the subtribe Carduinae (tribe Cardueae, Compositae). -Englera 21.
Holmgren, P. K. & Holmgren, N. H. 1998+ (continuously updated): Index herbariorum: a global directory of public herbaria and associated staff. -Published on the Internet at http://sweet gum.nybg.org/ih/
Huber-Morath, A. 1975: Cousinia in P. H. Davis (ed.), Flora of Turkey and the East Aegean Islands vol.. 5: 329-353. –Ediburgh University Press.
Knapp, H. D. 1987: On the distribution of the genus Cousinia (Compositae). -Pl. Syst. Evol. 155: 15-25.
Koch, C. 1851: Beiträge zu einer Flora des Orientes. - Linnaea 24: 305-624.
Lopez-Vinyallonga, S., Mehregan, I., Garcia-Jacas, N., Tscherneva, O. Susanna, A. and Kadereit, J. W. 2009: Phylogeny and evolution of the Arctium-Cousinia complex (Compositae, Cardueae-Carduinae). – Taxon 58:153-171.
Mabberley, D. J. 1990: The plant book. -Cambridge, UK: Cambridge University Press.
Mehregan, I. 2008: Systematics, phylogeny and biogeography of Cousinia-Asteraceae (Ph.D. thesis).?
Mehregan, I & Kadereit, J. W. 2008: Taxonomic revision of Cousinia sect. Cynaroideae (Asteraceae, Cardueae). Wildenowia 38: 293-362.
Rechinger, K. H. 1972: Compositae-Cynareae I. Cousini in K. H. Rechinger (ed.). Flora Iranica, no. 90. –Graz, Austria.
Rechinger, K. H. 1979: Compositae III-Cynareae. Cousinia in K. H. Rechinger (ed.). Flora Iranica, no. 139A: 108-153. –Graz, Austria.
Rechinger, K. H. 1986: Cousinia: morphology, taxonomy, distribution and phytogeographical implication. -Proceedings of the Royal Society of Edinburgh 89 B: 45-58.
Susanna, A., Garcia-Jacas, N., Vilatersana, R. & Garnatje, T. 2003a: Generic boundaries and evolution of characters in the Arctium group: a nuclear and chloroplast DNA analysis. -Collectanea Botanica, Barcelona 26: 101-118.
Susanna, A., Garcia-Jacas, N., Vilatersana, R. Garnatje, T., Valles, J. & Ghaffari, S. M. 2003b: New chromosome counts in the genus Cousinia and the related genus Schmalhausenia (Asteraceae, Cardueae). –Bot. J. Linn. Soc. 43: 411-418.
Susanna, A. & Garcia-Jacas, N. 2006 [“2007”]: Tribe Cardueae Cass. (1819). – Pp. 123-147 in Kadereit J. W. & Jeffrey C. (volume ed.), the families and genera of vascular plants 8. – Berlin, etc.
Tscherneva, O. V. [Cherneva O. V.] 1962: Compositae-Cousinia in Shishkin B. K. & Bobrov E. G. (ed.), Flora SSSR 27: 108-357. - Moscow and Leningrad [English translation: Enfield. 1993].
Tscherneva, O. V. 1985: Chromosome number in the species of the genus Cousinia (Asteraceae) in the flora of the USSR. -Botaniceskij Zurnal 70: 856-857 (in Russian).
Winkler, C. 1892: Synopsis specierum generic Cousinia Cass. -Acta Horti Peteropolitanis 12 (2): 181-286.
Winkler, C. 1897: Mantissa synopsis specierum generic Cousinia Cass. -Acta Horti Peteroplitanis 14 (2): 187-243.