Afzal-Rafii, Z. 1980: Contribution to Cytotaxonomy of Cousinia in Iran (in French). – Biol. Ecol. Medit, 7: 9-14.
Attar, F. & Ghahreman, A. 2000: Two new species and a new record of the genus CousiniaCass., Cynaroideae (Asteraceae) from Iran. -Iran J. Bot, 8: 259-269.
Attar, F. & Ghahreman, A. 2002: New taxa of genus of Cousinia. -Iran J. Bot, 92: 161- 169.
Attar, F., Ghahreman, A., Saber, A. & Zarre, S. 2005: A new species of the genus Cousinia (Compositae, sect. Stenocephala) from SW of Iran. - Iran J. Bot, 11 (1): 65-69.
Ghaffari, S. M. & Djavadi, S. B. 1998: Chromosome studies and distribution of nine species of Cousiniasection Stenocephalae (Asteraceae) in Iran. –Bul. de la Soci. Neuchateloise des Sci. Natur, 121: 61-68.
Ghaffari, S. M., Attar, F. & Ghahreman, A. 2000: Distribution and chromosome studies on some species of Cousinia Cass. (section Cyanoideae) from Iran. –Pakistan J. Bot, 32: 311-316.
Djavadi, S. B. 2005: New or rare chromosome counts in ten species of Cousinia from Iran. –Rostaniha,6: 61-70.
Djavadi, S. B. 2007: Cousinia papilosa, a new species from eastern Iran including chromosome count and palynological studies. –Rostaniha, 8: 63-73.
Hansen, H. V. 1991: SEM-studies and general comments on pollen in tribe Mutisieae (Compositae) sensu Cabrera. -Nord. J. Bot,10: 607-623.
López-Vinyallonga, S.,Mehregan, I., Garcia-Jacas, N., Tscherneva, O., Susanna, A. & Kadereit, J. W. 2009. Phylogeny and evolution of the Arctium-Cousiniacomplex (Compositae, Cardueae-Carduinae). –Taxon 58 (1): 153-171.
Lundberg, J. 2009: Asteraceae and relationships within Asterales. -In: Funk, V. A., Susanna, A., Stuessy, T. F. & Bayer, R. J. (eds.), Systematics, Evolution, and Biogeography of Compositae, Pp. 157-169. IAPT, Vienna, Austria.
Mehregan, I. & Kadereit, J. W. 2008. Taxonomic revision of Cousiniasect. Cynaroideae(Asteraceae, Cardueae). -Willdenowia38: 293-362.
Meo, A. A. M. & Khan A. 2003: Palynological Observations on The Genera Zoegia, Cousiniaand Carduus in the tribe Cynaroideae (Compositae) From Pakistan. –Pak. J. Weed Sci. Res, 9: 243-252.
Podani, J. 2000: Introduction to the Exploration of Multivariate Biological Data. -Backhuys Publisher, Leiden, Pp 407.
Rechinger K. H. 1970: Flora Iranica 90 (Cynerae), -Akademische Druck-U. Verlagsanstalt, Graz, Pp. 87-95.
Scotland, R. W., Olmstead, R. G. & Bennett, J. R. 2003: Phylogeny reconstruction: the role of morphology. -Syst. Biol, 52: 539-548.
Sheidai, M., Mehdigholi, K., Ghahreman, A. & Attar, F. 2006a: Cytogenetic study of the genus Cousinia(Asteraceae, section Serratuloideae) in Iran. –Genet. Mol. Bio, 29: 117-12.
Sheidai, M., Mahdigholi, K., Ghahreman, A. & Attar F. 2006b: Cyto-morphological study of the genus Cousinia sect. Serratuloideae in Iran. -The International Compositae Alliance: A Symposium: Botanic Institute of Barcelona., Barcelona, Spain, 3-9 July. Pp. 47.
Urtubey, E. & Telleria, M. C. 1998: Pollen morphology of the subfamily Barnadesioideae (Asteraceae) and its phylogenetic and taxonomic significance. -Rev. Palaeobot. Palynol, 104: 19-37.
Wortley, A. H., Funk, V. A., Robinson, H., Skvarla, J. J. & Blackmore, S. 2007: A search for pollen morphological synapomorphies to classify rogue genera in Compositae (Asteraceae). -Rev. Palaeobot. Palynol, 146: 169-181.
Zafar, M., Ahmad, M. & Khan, M. A. 2007: Palynology of Family Asteraceae from Flora of Rawalpindi-Pakistan. -Inte. J. Agri. Biol., 9: 156-161.