Assadi, M. 2009: Four new species of Cousinia Cass. (Asteraceae) from Iran. –Iranian J. Botany 15 (1): 36-44.
Assadi, M. 2011: Two new species from the Genus Cousinia (Asteraceae) from Iran. –Iranian J. Botany 17 (1): 6-9.
López-Vinyallonga, S., Mehregan, I., Garcia-Jacas, N., Tscherneva, O., Susanna, A. & Kadereit, J. W. 2009: Phylogeny and evolution of the Arctium-Cousinia complex (Compositae, Cardueae-Carduinae). – Taxon 58:153–171.
Mehregan, I. & Kadereit, J. W. 2008: Taxonomic revision of Cousinia sect. Cynaroideae (Asteraceae, Cardueae). –Willdenowia 38: 293-362.
Mehregan, I. & Kadereit, J. W. 2009: The role of hybridization in the evolution of Cousinia s.s. (Asteraceae). – Willdenowia 39: 35-47.
Mehregan, I., Assadi, M. & Attar, F. 2003: Cousinia (sect. Haussknechtianae) gatchsaranica, a new species from SW Iran. – Willdenowia 33: 107-111.
Mehregan, I., Djavadi, S. B. & Pahlevani, A. H. 2010: Cousinia (sect. Haussknechtianae) karkasensis, a new species from Karkas Mts. in central Iran. -Iranian J. Bot. 16 (2): 200-203.
Rechinger, K. H. 1972: Compositae-Cynareae1. Cousinia in Rechinger, K. H. (ed.), Flora Iranica 90. – Graz, Austria.
Rechinger, K. H. 1979: Compositae III-Cynareae. Cousinia in Rechinger, K. H. (ed.), Flora Iranica 139a: 108-153 – Graz, Austria.
Rechinger, K. H. 1986: Cousinia: morphology, taxonomy, distribution and phytogeographical implication. – Proceedings of the Royal Society of Edinburgh 89 B: 45-58.
Susanna, A., Garcia-Jacas, N., Vilatersana, R. & Garnatje, T. 2003: Generic boundaries and evolution of characters in the Arctium group: a nuclear and chloroplast DNA analysis. – Collect. Bot. (Barcelona) 26:101-118.
Susanna, A. & Garcia-Jacas, N. 2006 [“2007”]: Tribe Cardueae Cass. in Kadereit, J. W. & Jeffrey, C. (volume ed.), The families and genera of vascular plants 8: 123-147. –Berlin.
Thiers B. 2008+ [continuously updated]: Index herbariorum: A global directory of public herbaria and associated staff. – http://sweetgum.nybg.org/ih/.